Baena & Torres, 2013b (ecology); Heiss, 1979a (fig.), 1994 (types). Distribution: Carapezza, 2002a (Jordan); Gadalla, 2004 (Sinai); Heiss, 2006 (Albania); Linnavuori, 2011 (Iran); Baena & Torres, 2013b (Andorra Balearic Islands Portugal Spain); Parmain et al., 2013 (Corsica France); Carapezza & Mifsud, 2015 (Malta); Heckmann et al., 2015 (Crete).