Cobben, 1958b; Dethier & Chérot, 1997; Southwood & Woodroffe, 1957; Wagner, 1960l.
Distribution: Misja, 1973 (Albania); Protić, 1998a (Serbia); Drapolyuk, 2001 (Georgia); Konstantinov & Namyatova, 2009 (Mongolia); Khaghaninia et al., 2010a, 2010c (Iran); Baymak & Kıyak, 2022 (Kosovo).