Hradil et al., 2007 (biol., distr.); Parshad, 1957a (chrom.).
Distribution: Protić, 2001a (Montenegro Serbia); Pettersson & Coulianos, 2004 (Sweden); Ollikainen & Rinne, 2005 (Finland); Hradil et al., 2007 (Arab Emirates Switzerland); Ghahari et al., 2012c (Iran); Skipper & Tolsgaard, 2013 (Denmark); Heckmann et al., 2015 (Crete); Frey et al., 2016 (Switzerland); Carapezza et al., 2017 (Arab Emirates); Baymak & Kıyak, 2022 (Kosovo).