Grozeva, 1995 (karyotype, male reproductive system); Henry & Froeschner, 2000 (suppl. World cat.); V.G. Putshkov, 1974a (biol.).
Distribution: Protić, 2001a (Serbia); Heckmann et al., 2015 (Crete); Gogala et al., 2020 (Slovenia); Baymak & Kıyak, 2022 (Kosovo).