Grozeva, 1995 (karyotype, male reproductive system); V.G. Putshkov, 1974a (biol.); Schwoerbel, 1956 (biol.).
Distribution: Protić, 2001a (Montenegro Serbia); Kment, 2005 (Czech Republic); Şerban, 2010 (European Turkey); Heckmann et al., 2015 (Crete); Matocq & Azar, 2023 (Lebanon); Baymak & Kıyak, 2022 (Kosovo).