Roubal, 1959a (maurum and brevicolle); Zheng & Zou, 1981a.
Distribution: Roubal, 1959a (Albania); Ashlock, 1977 (Canada); El-Moursy et al., 2001 (Sinai); Protić, 2001a (Montenegro Serbia); Linnavuori et al., 2014b (Arab Emirates); Heckmann et al., 2015 (Crete); Gossner et a., 2017 (Germany).