Rouault et al., 2005 (biol.); Seabra, 1930b.
Distribution: Gessé et al., 1994 (Andorra); Protić, 2001a (Serbia); Protić & Roganović, 2004 (Montenegro); Rouault et al., 2005 (China: SW); Skipper, 2009 (Denmark); Korcz, 2010 (Poland); Heckmann et al., 2015 (Crete); Aukema (Azores!); ZMAN (Canary Islands!).