Andersen, 1994 (syst.); Damgaard, 2006, 2008a (phyl.); Klingenberg, 1992 (geogr. var.); Wagner & Zimmermann, 1955 (syst.). Distribution: Protić, 1998a (Montenegro Serbia, as costae costae); Saleh Ahmed & Gadalla, 2005b (Egypt); Ghahari et al., 2013a (Iran).