Heiss, 1998a; Morkel, 2017 (ecology); Štys, 1974 (morph.); Vásárhelyi, 1980a (larvae).
Distribution: Protić, 2001a (Bosnia Hercegovina Montenegro Serbia); Heiss, 2006 (Romania); Heiss & Péricart, 2007 (Syria Turkmenistan Uzbekistan); Parmain et al., 2013 (France); Baymak & Kıyak, 2022 (Kosovo).