Carapezza, 1997; Kaplin, 1993 (biol.); Linnavuori, 1986a; V.G. Putshkov, 1976b, 1976d (biol.); Wagner, 1964e (Itupona). Distribution: Kerzhner & Josifov, 1999 (Israel); Carapezza, 2002a (Jordan); Konstantinov & Namyatova, 2008 (Asian Kazakhstan Uzbekistan); Ghahari & Chérot, 2014 (Iran); Linnavuori et al., 2014a (Arab Emirates).