Schuh, 2001 (male genitalia); Wagner, 1967a.
Distribution: Protić, 1998a (Serbia); Aukema et al., 2002 (Belgium); Carapezza, 2002a (Jordan); Heiss & Linnavuori, 2002 (Iran); Ghahari & Chérot, 2014 (Iran); Heckmann et al., 2015 (Crete); Baymak & Kıyak, 2022 (Kosovo).