Kelton, 1982a; Wyniger & Burckhardt, 2003 (male genitalia).
Distribution
Protić, 1998a (Montenegro Serbia); Aukema et al., 2002 (Belgium); Coulianos, 2003 (Estonia); A. Gogala, 2006 (Slovenia); ); Linnavuori, 2007a (Iran); Konstantinov & Namyatova, 2009 (Armenia Kirgizia Uzbekistan); Söderman & Rintala, 2009 (Lithuania); Konstantinov & Vinokurov, 2011 (China NW); Ghahari & Chérot, 2014 (Iran).