Josifov, 1959b; Kment et al., 2006a (biol., distr.).Sienkiewicz, 1963; Wagner, 1968t.
Distribution: Protić, 1998a (Serbia); Kment et al., 2006a (Slovakia Switzerland); Vinokurov & Golub, 2007 (Russia: WS); Konstantinov & Zinovyeva, 2017 (Russia: CT ST); Baymak & Kıyak, 2022 (Kosovo).