P.V. Putshkov, 1988b (key), 1994b (monogr.). Distribution: Coulianos, 2003 (Estonia); Kment et al., 2003 (Byelorussia Czech Republic); Tolsgaard, 2005 (Denmark); Strauß & Günther, 2006 (Germany); Hradil et al., 2008 (Slovakia); P.V. Putshkov & Moulet, 2010 (Kirgizia); Rintala & Rinne, 2010 (Finland);; Foster, 2013 (Great Britain); Frieß et al. 2013 (Austria); Ghahari et al., 2013b (Iran).