Dispons, 1955a (biol.), 1964e (key, map), 1965b (key, tax.), 1965c (tax.), 1968a (fig., tax.); Poisson, 1933b (eggs); Ribes, 1961a (fig., key); Tamanini, 1981 (habitat).
Distribution: Protić, 2010a (Montenegro); Moulet, 2004 (Iran); Whitehead, 2006 (Great Britain); P.V. Putshkov & Moulet, 2010 (Asian Turkey Iraq Saudi Arabia Syria Yemen: Socotra); Heckmann et al., 2015 (Crete).