Boulard & Coffin, 1991 (biol.); Jaczewski, 1934a (syn., tax.).; Kormilev, 1971a, 1973 (distr., tax.); Rieger, 1976 (morph.); Takahashi, 1921, 1923 (biol.).
Distribution: Katbeh et al., 2000 (Jordan); Linnavuori et al., 2011 (Arab Emirates); Polhemus & Polhemus, 2012 (Israel); Ghahari et al., 2013a (Iran); Kment et al., 2020 (Oman).